My markers make most of pens smudge (and I found out only after I had most of this done that my copic pens don’t smudge), so I tried lightly penciling everything, colored over them, then ink over that, but that caused a lot of mistakes - The pencils wouldn’t erase after the marker colored over them, and in some cases, I couldn’t see where the lines were supposed to be. Ended up making a lot of corrections digitally.
Oh well. Chalk it up as learning experience.